Perceptual Oscillation:
A few years ago it would not have been considered appropriate by the
leaders in the field to use the word “oscillation” to describe the
irregular, variable and often-voluntary alternations of perceptual
rivalry. The timing of rivalry was usually ignored, or even disguised
by the “phase normalisation” process where individual variations in
rivalry rate were factored out.
That has changed.
There is now recognition that there may be an underlying oscillatory
process that forms a basis for the variety of perceptual rivalries.
There is no agreement on the location of the oscillator, with
Hugh Wilson’s “g” factor of cortical networks at one extreme and my own
suggestion of a sub-cortical, interhemispheric oscillator at the other.
Here are some of the new facts that have made it acceptable to talk
about rivalry as a perceptual oscillation (numbers in brackets refer to
my publication list):-
1. The pattern of oscillations can be predictably changed by caloric
vestibular stimulation (157) and TMS (157,177). Since there are 2
perceptual alternatives and 2 hemispheres, there are 4 possible effects
of the manipulations, but only the predicted one is seen. It is easier
to manipulate the Left hemisphere than the Right, but in later
experiments we succeeded in producing clear effects from Right
hemisphere TMS also (177).
2. The frequency of rivalry oscillations is coupled in the same
individual across different forms of rivalry. For example, a “fast
switcher” on binocular rivalry will also be a “fast switcher” on Plaid
Motion Rivalry (190). This is true despite the wide inter-individual
variation in rivalry rate.
3. Motion-induced Blindness is a form of rivalry, even though it does
not show all of the accepted outward attributes, such as the
symmetrical pair of alternative percepts that allows one to guess which
percepts will alternate. The identity of MIB with rivalry was shown
using neuropharmacology (LSD produces identical changes in the timing
of both binocular rivalry and MIB), a demonstration that Levelt’s
second proposition also applies to to MIB and the frequency coupling
phenomenon (175).
4. Rivalry timing has a neural mechanism that uses 5HT1a receptors, not
the 5HT2a receptors that are prominent in visual cortex. This finding
shifts the focus on the rivalry timer to the brainstem where 5HT1a
receptors predominate (183).
5. Although rivalry rate is very stable in one individual, rare
individuals with extensive training in meditation, can change their
rivalry rate (189).
6. Rivalry oscillations cease (or become too fast to be noticeable)
during laughter (186)